in the fused structure of the pygidium, intersegmental lines are usually lost. However, the segmentation can be deduced from the positions of marginal structures, e.g. abdominal segment 8 bears the median lobes, segment 7 bears the second lobes, segment 6 bears the third lobes and segment 5 bears the fourth lobes. Each pygidial segment also bears a dorsal and ventral marginal seta on each side