Douglas, 1887
Appearance in life
Scale cover of adult female narrow and very elongate, almost parallel-sided, flat to moderately convex, bright brown to black, with terminal exuviae. Males have not been documented.
Morphology
Adult female not pupillarial; body elongate, usually slightly fusiform; membranous except for the pygidium; lateral lobes of free abdominal segments slightly developed, marginal sclerotized spurs absent. Pygidium with reticulated pattern on median and submedian areas around anus, reaching to fourth segment. Two pairs of pygidial lobes present: median lobes well separated, not zygotic; often rounded, with finely serrated edges; a pair of marginal setae present between them. Second lobes much smaller than median lobes. Ventral sclerotizations and paraphyses present, short ones associated with the median lobe, and second lobe with slender ventral paraphyses extending anteriorly from base of inner lobule. Gland spines well developed, present on pygidial and at least some prepygidial abdominal segments; absent from between the median lobes; replaced by duct tubercles when anterior to abdominal segment II. Anus oval to circular, situated slightly posterior to centre of pygidium; vulva situated near centre of pygidium. Marginal macroducts usually numbering 2 on either side of the pygidium, one between median and second lobes and one just lateral to second lobe; never present between median lobes; each marginal macroduct enlarged, with a thick sclerotized rim and with the long axis of the orifice perpendicular to the pygidial margin. Dorsal microducts sparse, present on abdominal segments only. Ventral microducts numerous on abdominal segments and extending as far forward as antennal tubercles. Submarginal dorsal bosses present. Duct tubercles may be present on the prepygidium as far forward as the mesothorax. Perivulvar pores, if present, rather few and arranged in 5 groups. Stigmatic disc pores present by anterior spiracles only. Antennae each with 2 or more setae.