MacGillivray, 1921
Appearance in life
Scale cover of adult female oval or subcircular, slightly convex, white, with yellow exuviae usually subcentral to almost marginal. Males not recorded.
Morphology
Adult female not pupillarial; body oval to pyriform, the anterior edge becoming sclerotized with maturity; pygidium also partly sclerotized; lateral lobes of free abdominal segments only slightly developed, never with marginal sclerotized spurs. Pygidium shallow and rounded, with a slight apical notch formed by divergent apical lobes. Median lobes well developed, divergent, separate, never zygotic; each lobe asymmetrical, with a long, serrate inner margin and a short out one; a pair of marginal setae present between them. Second and third lobes similar in size, smaller than median lobes, bilobed; fourth lobes bilobed but less well developed normally reduced to a single poorly defined lobe. Ventral paraphyses usually present, arising from basal corners of median lobes at least. Gland spines present: on pygidium, long, present singly, each fimbriate and containing 2 microducts, including a pair between the median lobes that are shorter than the lobes: gland spines on prepygidial segments simple, with single ducts. Anus oval, situated more or less at the centre of pygidium; vulva also situated at centre of pygidium. Marginal macroducts numbering 6 on either side (1 between median and second lobes, 2 associated with the third lobe, 2 on the fifth segment and 1 on a slight spur on the fourth segment); 1 also present between median lobes; each marginal macroduct of moderate size, with a sclerotized rim and with the long axis of the orifice perpendicular to the pygidial margin. Dorsal macroducts of two types; several ducts the same size as marginal ducts, present submarginally; a second type, smaller than a marginal duct, scattered in submarginal and submedian zones on pygidium and prepygidial segments. Submarginal dorsal bosses absent. Pointed ventral duct tubercles present on some prepygidial segments. Perivulvar pores absent. Stigmatic disc pores few, 1-3 present by each anterior spiracle. Antenna usually with only 1 seta, curved.